Knuckles Biographical information Home Earth Occupation Criminal Physical description Species Human Gender Male Out of universe information Era(s) 1987 TV series First appearance Leonardo Lightens Up Created by Dan DiStefano Voiced by Rob Paulsen Teachers and Students [Source]Knuckles is a character in the 1987-1996 animated TV series. He appears in the episode Leonardo Lightens Up, and is one of G. Cleff's henchmen. References ↑ Wildcat 7812 (9 December 2013). List of classic TMNT characters (English). Deviant Art. Retrieved on May 14, 2017. Retrieved from "https://turtlepedia.fandom.com/wiki/Knuckles?oldid=543688" Categories: Male characters Humans 1987 TV series characters Antagonists (1987 TV series) Musicians (in-universe) Community content is available under CC-BY-SA unless otherwise noted.